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Eocetus

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Eocetus
Temporal range: Late Eocene, 40.4–37.2 Ma
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Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Artiodactyla
Infraorder: Cetacea
Family: †Protocetidae
Subfamily: †Georgiacetinae
Genus: †Eocetus
Fraas, 1904
Type species
†Eocetus schweinfurthi
Other species
Synonyms

Eocetus is an extinct protocetid early whale known from the early-late Eocene (Bartonian, 40.4 to 37.2 million years ago) Giushi Formation in Gebel Mokattam, (30°00′N 31°18′E / 30.0°N 31.3°E / 30.0; 31.3, paleocoordinates 24°54′N 26°48′E / 24.9°N 26.8°E / 24.9; 26.8) outside Cairo, Egypt.[1][2] Fossil remains have also been discovered in the Aridal Formation of the Sahara Desert in southwestern Morocco.[3]

The type specimen was first named by Fraas as Mesocetus schweinfurthi. However, the name Mesocetus was previously used causing a change to the species name to Eocetus schweinfurthi. Since the genus was first described in the early 20th century, several other specimens, mostly isolated vertebrae, have been attributed to Eocetus, but the taxonomic status of these widely distributed specimens remain disputed.

Discovery and taxonomy

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Fraas 1904a described "Mesocetus schweinfurthi" based on a dorsoventrally compressed skull with only I2 in situ, a specimen supposedly originating from a 40 Ma Tethyan deposit at Mokattam. Fraas also referred two isolated teeth, P4 and M1, to the skull and the most important of his specimens is not the deformed skull, but the upper molar which retains three roots and a worn but well-developed protocone. Georg August Schweinfurth, a German palaeontologist who explored Mokattam in the 1880s, mentioned the quarriers there very eagerly offered "shark teeth" to tourists and that scientists and fossil collectors regularly bought their specimens from this source. There is reason to assume that Fraas were among them and that at least his two isolated teeth were described without direct knowledge of their original locality and stratigraphic context. Notwithstanding that the stratigraphic information supplied by Fraas and his contemporaries can be difficult to interpret, the geology of Egypt is well studied, and both the skull and the accompanying teeth are most likely Bartonian in age—older and significantly more primitive than any other cetacean specimen known from Egypt at that time.[4][5] Fraas soon discovered that the name "Mesocetus" was already occupied, and changed the name of his "Urform Protocetus" to Eocetus.[6]

Fraas also attributed two isolated vertebrae to his new genus, both of which Uhen 1998 moved to Basilosaurus drazindai. A holotype, described by Stromer 1903 as Zeuglodon macrospondylus, was discovered in Egypt. It is later used for comparison for other vertebrae. Stromer 1908 attributed two other vertebrae from Mokattam to Eocetus. These two vertebrae were lost for many years until Uhen 1999 described two bones that he discovered in a museum in Germany and appeared to fit Stromer's description. Uhen based his assignment of his own North American genus (see below) to Eocetus on the similarities to Stromer's vertebrae.[7]

Previously assigned specimens and taxa

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Uhen 1999 described a new species, Eocetus wardii, from the late Lutetian (~42 Ma) of North Carolina based on more complete material: a partial skull, a few thoracic, lumbar and caudal vertebrae, ribs and an innominate fragment. Uhen initially argued that the innominate would have been sufficiently large to support a weight-bearing hind limb—suggesting the animal was a protocetid, a group of more primitive archaeocetes—but also has anatomical features in common with basilosaurids—more derived and fully aquatic archaeocetes. Uhen also noted that the composition of the ribs and vertebrae is different from that of other archaeocetes and sirenians: a core of light trabecular bone is surrounded by layers of dense cortical bone.[8] Due to this mosaic of protocetid and basilosaurid features, Geisler, Sanders & Luo 2005 regarded Uhen's specimen as unique among North American archaeocetes and an interesting find, but questioned the validity of Uhen's attribution (including that of Stromer's two vertebrae). This suspicion was confirmed by Gol'din & Zvonok 2013, who reassigned E. wardii remains to the genus Basilotritus (and hence Basilosauridae),[9] while Gol'din have concluded that the original interpretation of its innominate were incorrect, and that the hips could have been more reduced and Basilosaurus-like than Uhen first thought.[10]

Uhen & Berndt 2008 described another partial lumbar vertebra discovered in Rohrdorf, Bavaria, Germany, in 2003. They attributed it to Eocetus sp.—the first confirmed protocetid from Europe—and argued that this specimen further supports the hypothesis that protocetids were aquatic to the extent that they managed to spread over the world.[11][12]

Two vertebrae, a thoracic and a lumbar, discovered on a riverbed in Virginia in 2009 were referred to "Eocetus" wardi by Weems et al. 2011.[13][14]

Uhen et al. 2011 described a still unnamed Bartonian protocetid from Peru based on the posterior portion of a skull, seven partial vertebrae, and ribs from and adult individual. Uhen et al. considered this specimen closely related to Eocetus based on vertebral morphology.[15]

Goldin, Zvonok & Krakhmalnaya 2012 described two vertebra, a thoracic and a lumbar, from a subadult individual found in Ukraine. They considered them comparable to those Uhen described in 1999 and attributed their specimen to "E." wardii.[16]

References

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Notes

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  1. ^ Eocetus in the Paleobiology Database. Retrieved 1 July 2013.
  2. ^ Gebel Mokattam, upper (Eocene of Egypt) in the Paleobiology Database. Retrieved 1 July 2013.
  3. ^ Gingerich, Philip D.; Zouhri, Samir (November 2015). "New fauna of archaeocete whales (Mammalia, Cetacea) from the Bartonian middle Eocene of southern Morocco". Journal of African Earth Sciences. 111: 273–286. doi:10.1016/j.jafrearsci.2015.08.006.
  4. ^ Bianucci & Gingerich 2011, p. 1174
  5. ^ Gingerich 1992, pp. 71–2
  6. ^ Fraas 1904b, p. 374
  7. ^ Geisler, Sanders & Luo 2005, p. 47
  8. ^ Uhen 1999, Abstract
  9. ^ Gol'din & Zvonok 2013, Abstract
  10. ^ Gol'din, P. (2014). "Naming an Innominate: Pelvis and Hindlimbs of Miocene Whales Give an Insight into Evolution and Homology of Cetacean Pelvic Girdle". Evolutionary Biology 41: p. 473—479. doi:10.1007/s11692-014-9281-8
  11. ^ "Eocetus sp". Collections Search Center, Smithsonian Institution. Retrieved 1 July 2013.
  12. ^ Uhen & Berndt 2008, Abstract
  13. ^ Weems et al. 2011, Abstract
  14. ^ Weems et al. 2011, Material, p. 273
  15. ^ Uhen et al. 2011
  16. ^ Goldin, Zvonok & Krakhmalnaya 2012, Introduction

Sources

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